The impact of livestock grazing on desert vegetation in Khirthar National Park, Pakistan, was investigated by comparing dry and wet season plant species composition, richness, cover, and a grazing index for quadrats outside (‘‘open’’) and inside (‘‘exclosed’’) native mammal breeding enclosures that had excluded livestock for 6 years. A total of 93 plant species were recorded in the dry season, 88 species in exclosed quadrats and 50 in open quadrats. While only 5 species were unique to open quadrats, 43 species were found only in the exclosed quadrats. Species richness was higher in the exclosures because of the presence of more grass and herb species, while grazing was higher in the open. After rain, species richness and cover were significantly higher than in the dry season because of the growth of summer ephemeral herbs and grasses, but richness was no longer different between the exclosure and open treatments. Although some herbaceous species may have been adversely affected by livestock grazing, overall species richness suggests strong ecosystem resilience to grazing, with levels no different after seasonal rains regardless of grazing level. Many grass and herb species absent from open sites during the dry season reappeared after rain, which suggests that livestock grazing may eliminate them as the dry season proceeds, but that a soil seed or bud bank persists. 

In and around the Great Basin, United States, restoration of shrub steppe vegetation is needed where rangelands are transitioning to annual grasslands. Mechanical seedbed preparation can aid native species recovery by reducing annual grass competition. This study was designed to investigate the nature and persistence of hydrologic and erosion impacts caused by different mechanical rangeland seeding treatments and to identify interactions between such impacts and related soil and vegetation properties. A cheatgrass (Bromus tectorum L.)-dominated site was burned and seeded with native grasses and shrubs in the fall of the year. An Amazon-drill and a disk-chain seeder were used to provide varying levels of surface soil disturbance. An undisturbed broadcast seeding was used as a control. Simulated rainfall was applied to 6 large (32.5-m2) plots per treatment over 3 growing seasons at a rate of 63.5 mm h-1. Rainfall was applied for 60 minutes under dry antecedent moisture conditions and for 30 minutes, 24 hours later under wet antecedent moisture conditions. The disk-chain created the largest reduction in infiltration and increase in sediment yield, which lasted for 3 growing seasons posttreatment. The Amazon-drill had a lesser impact, which was insignificant after the second growing season posttreatment. Surface soil properties showed little correlation with treatment-induced hydrologic and erosion impacts. Hydrologic recovery was strongly correlated with litter dynamics. The seeding treatments were unsuccessful at establishing seeded plant species, and the site once again became dominated by cheatgrass. A continuous upward trend in biomass production and surface litter cover was observed for all treatments between the beginning and end of the study because of cheatgrass invasion. Although the initial goal of using mechanical seeding treatments to enhance recovery of native grass species failed, cheatgrass production provided sufficient biomass to rapidly replenish surface litter cover necessary for rapid hydrologic stability of the site. 

Soil nutrient availability influences plant invasions. Resin capsules were used to examine soil nutrient bioavailability along 2 sagebrush-grassland elevation transects in the east Tintic Range (Utah) and Shoshone Range (Nevada). In the fall of 2001, treatments were applied to 3 replicate plots at each site, which included prescribed burning, herbaceous vegetation removal, and controls. Cheatgrass (Bromus tectorum L.) was overseeded in small subplots within each treatment. Following treatments in each plot, resin capsules were installed at 15-cm depth in a shrub interspace and a B. tectorum-overseeded area. Nutrient availability was integrated during late fall to spring and spring to late fall for 2 years. Herbaceous vegetation removal increased availability of nitrate (Nevada and Utah) and Ca and Mg (Nevada only) but only during the second sampling period (growing season). Availability of K and ortho-P (Nevada and Utah) and nitrate (Nevada only) was greater on prescribed burned plots. For Utah, availability of ortho-P, K, Ca, Mg, and Fe generally increased with increasing elevation. Availability of Ca, Mg, K, and Fe was greatest during late fall to spring integration periods for Nevada. Overseeding with B. tectorum interacted with the burn treatment to influence availability of Ca, Mg, and Fe (Nevada sites only). Patterns of nutrient availability can be explained by a combination of decreased root uptake in relation to mineralization, differences in soil water content with season and elevation, and nutrient release from vegetation and soil as a consequence of prescribed burning. Herbaceous vegetation removal and burning can raise nitrate availability and increase risk of invasion by nitrophilic species such as B. tectorum. Nutrient availability can be out of phase with plant growth; plants capable of taking up nutrients during cold periods may have a competitive advantage. Resin capsules have utility in quantifying the effects of treatments on the availability of many soil nutrients. 

A field study was conducted to investigate the effects of existing adult neighbors and gap size on the process of seedling establishment in Leymus chinensis. Seeds of L. chinensis were added to artificially created gaps in a degraded steppe in North China. Neighbor root exclusion was accomplished using polyvinyl chloride tubes sunk in the soil of gaps. Seedling emergence and survivorship was greater in gaps than in control areas, but growth performance was higher only in larger gaps (20 cm and 40 cm in diameter) with neighboring roots present and in all gaps without neighboring roots present. Seedlings produced no tillers in the control and few tillers in 10-cm- and 20-cm-diameter gaps. Seedlings had more tillers in the largest gaps (40 cm) without root exclusion and in all sizes of gaps with root exclusion. Differences between gaps in light levels can explain the patterns of emergence. However, root exclusion was the major factor that increased seedling growth performance. These results confirm that L. chinensis is a gap-enhanced species and suggest that restoration of degraded grassland needs to ensure that large light gaps and low belowground competition are regularly maintained to maximize successful seedling recruitment. 

Russian knapweed (Acroptilon repens [L.] DC.), an exotic perennial forb, has invaded many native ecosystems in western North America. Russian knapweed’s success is attributed to allelopathy, extensive tap rooting, zinc accumulation in soils, and a lack of North American predators. Revegetation following chemical control slows exotic reestablishment, but the impacts of Russian knapweed-invaded soils on the establishment of native forbs and shrubs have not been determined. In a greenhouse experiment, we monitored the establishment of two native forbs, Indian blanketflower (Gaillardia aristata Pursh) and purple prairie clover (Dalea purpurea Vent.) and two native shrubs, winterfat (Krascheninnikovia lanata [Pursh] A.D.J. Meeuse Smit syn. Ceratoides lanata) and Wyoming big sagebrush (Artemisia tridentata Nutt. subsp. wyomingensis [Hook.] Nutt.) in soils obtained from three Russian knapweed invasions and adjacent noninvaded areas. We analyzed soils collected near Greybull and Riverton, Wyoming, and Greeley, Colorado, for cation exchange capacity, organic matter, electroconductivity, pH, and total nitrogen, carbon, and plant-available potassium, zinc, manganese, copper, and phosphate. We documented seedling emergence of the four natives and Russian knapweed every two days for 14-17 weeks, harvested seedlings biweekly to assess their growth, and determined their zinc accumulation. All species established in invaded soil and seedlings were larger in invaded than in noninvaded soils. Invaded rangeland soils had greater organic matter (8.6% and 1.1% in invaded vs. 2.5% and 0.4% in noninvaded soils) and lower pH (7.4 in invaded versus 8.0 noninvaded soils). Zinc concentrations in invaded soils (from 0.15 to 6.56 mg kg-1) were not high enough to limit plant growth. Reports that Russian knapweed is a hyper-accumulator of zinc are not supported by our seedling data, which suggests that previously invaded soils may not limit native seedlings. 

The potential for winter supplementation to increase juniper intake by goats on rangelands in the Edwards Plateau region of Texas was assessed in two experiments. The first experiment evaluated the effect on juniper intake of either no supplementation (negative control) or supplementation with corn, alfalfa, or cottonseed meal fed at an isonitrogenous protein level of 1.5 g kg body weight-1 for 12 days. Redberry juniper (Juniperus pinchotii Sudw.) consumption by individually penned Spanish, Boer 3 Spanish, Spanish 3 Angora, and Angora goats was measured on days 11 and 12. Each goat received each supplement in a complete 4 X 4 Latin square design. Juniper intake increased for goats supplemented with alfalfa and cottonseed meal (P = 0.001), but not for those supplemented with corn (P = 0.944). Boer 3 Spanish goats did not differ in levels of consumption (P = 0.085) from the other breeds. A second study investigated the effect of either no supplementation or soybean meal supplementation on juniper consumption by free grazing Angora and Boer 3 Spanish goats. Forty goats were assigned to four pasture groups by breed and previous juniper intake, and randomly allocated to either the treatment (supplementation) or control (no supplementation) regime in a complete block design. After 4 days of grazing and supplementation, fecal samples were collected to estimate percent of juniper in the diet using near-infrared spectroscopy. Goats were then rotated to another pasture. Juniper intake was highest for goats supplemented with soybean meal (P = 0.034). Breed of goat did not affect intake (P = 0.240). Goats previously categorized as high juniper consumers based upon prior measurements of juniper intake ate more juniper (P = 0.003) than those classified as low consumers. This research indicates that the effectiveness of goats for biological control of juniper can be improved with a high protein, low starch supplement. 

Plant invasions are rapidly becoming an important threat to the conservation of wildlands. Understanding how potentially invasive plants are dispersed to new habitats is a critical step in the process of understanding such invasions. Our objective was to characterize the potential for long-distance transport of plant species in the digestive tract of horses along recreational trails. We sampled horse dung along the first 4 000 m of the Lower Piney River trail in the White River Forest of western Colorado. We evaluated the seed content of each sample by applying standard methods for soil seed bank analysis. We found 20 species and 564 seedlings. Twelve of the species were graminoids, 6 were forbs, 1 was a shrub, and 1 was a tree. The species were evenly divided between natives and aliens, but 85% of the seedlings were aliens. An average of 47 seedlings emerged per sample, but the range was from 4 to 192. Our results make it clear that horses, and very likely all pack stock used on recreational trails, represent a potentially important dispersal vector for alien plants into western wildlands. 

We evaluated chukar (Alectoris chukar) watering patterns as well as habitat variables influencing water site selection in western Utah. Motion-sensing cameras and chukar dropping counts were primary techniques to evaluate watering patterns. We took vegetative and other habitat measurements at each water source (n = 43) to discriminate use from nonuse sites using logistic regression. Chukars watered during daylight hours with a modal hour from 1200 hours to 1300 hours daylight savings time. Annual patterns suggest limited use of water sources from November to May with first observed visits occurring in June and last observed visits in October. Shrub canopy cover was the only variable to discriminate between site types (P<0.01). Cross validation showed a predictive success rate of 84%. Significant differences were found between use and nonuse sites in terms of security cover (P < 0.01), but not total cover (P > 0.05). Chukars seem to have a loose shrub canopy threshold near 11% that is likely due to predation risk. Water sources meeting this threshold received use, whereas those not meeting this threshold did not. Increasing shrub canopy cover above 11% did not translate into increased water source use. Managers might want to consider annual patterns when setting hunt season timing and structure as well as judging sites for new water developments based on shrub canopy cover. More generally, these results suggest a behavioral constraint on the use of water sources as a function of predation risk—we should expect other species to demonstrate similar behavioral constraints. These constraints must be considered in any effort to determine benefits or impacts of water developments.